Appendix III: R1b Haplogroup Lineages

The principle R1b Haplogroup for Western Europe is M269, of which the following Sub-Haplogroups derive, with the main nations associated with them: U106 Germany; U198 England; S116 Spain & Portugal; U152 France, Northern & Central Italy; M529 Wales, Scotland & Ireland; M222 Scots-Irish.

M207 R

M479 R2 South Asia, Central Asia

M124 R2a

L263 R2a1

P267 R2a2

FGC21706 R2b / FGC50339 R2b1 

M173 R1 

M420 R1a 

M459 R1a1

M198 [M17] R1a1a

M417 R1a1a1

L664 R1a1a1a Northwest Europe

M458 R1a1a1b1a1 Northeastern Europe

Z280 [S466] R1a1a1b1a2 Finno-Ugric and Balto-Slavic speakers

Z282 R1a1a1b1a Eastern Europe, Russia

Z284 R1a1a1b1a3 Germanic, Scandinavia, Northern Ireland, Scotland, Northern England 

Z93 R1a1a1b2 Central Asia, Southwest Asia, South Asia, India

M343 / M415 R1b 

L278 R1b1

L754 R1b1a 

PH155 R1b1b Bahrain, Bhutan, Tajikistan, Turkey, Xinjiang & Yunnan, China

M335 R1b1b1 / PH200 R1b1b2 Anatolia, Yunnan, Ladakh, India

V88 R1b1a2 [R1b1c] Levant Arab/North Africa 

M18 R1b1a2a [R1b1c1] Sardinia, Lebanon

V35 [R1b1c2] Sardinia

V69 [R1b1c3] R1b1c4 Egypt 

L389 [L388] R1b1a1

P297 R1b1a1a

M73 / M478 R1b1a1a1 Central Asia 

M269 / M520 R1b1a1b [R1b1a2 2003-2005, R1b1c 2005-2008, R1b1b2 2008-2011, R1b1a1a2 2011-2020] Europe

L23 / S141 R1b1a1a2a [0] Caucasus, Southern Italy, Greece, Central Europe – Z2103 R1b1a1a2a2 Balkans, Turkey 

L51 / M412 / S167  R1b1a1a2a1 [1] Central France 

L11 [S127, L151] R-P310 [S128/S129] P311 [S128] R1b1a1a2a1a [2] Central England 

Austria – North Western Europe, Norway – R-Z18 R1b1a2a1a1b Scandinavia, Iceland, Low countries, British Isles – P107 [R1b1b2a1b] Germanic 

U106 / M405 / S21 [R1b1b2a1] R1b1a1a2a1a1 [3] Benelux, Netherlands, Frisia, England, Norway, Germany, Austria L257/S186 Germanic [L217, Z14, Z16, Z17, Z18, Z19]

R-S499 / Z301 R1b1a2a1c2 Z301>L48 Germanic >Z9 / S268>Z30>Z27 & Z2>Z7>CTS4099 (1) Sweden, Germany, UK, Ireland & >Z8> [Z338>Z11 / S274 (2) Scandinavia, Germany, UK, Ireland, Spain & FGC1954 (3) Netherlands, France, Britain, Poland & Z1>Z344 (4) Norway, Germany, UK & >Z346 / S512 (5) Germanic >Z343 /S387

Z381 / S263 Britain, Netherlands, Scandinavia, Germany, Poland. R-S264 / Z156 R1b1a2a1a1c1 – R-M1994 R1b1a2a1a1c3 

U198/ S29 / M467 [R1b1b2a1a] R1b1a1a2a1a1a [3a] Southern England and Northern Germany, Germanic (British Saxon) L1 / S26 [R1b1b2a1c] Southern & Eastern England, Norway, Southern Germany, Spain 

L48 / S162 Western Europe 

S116 / P312 R1b1a1a2a1a2[4] Spain and Portugal

Z196 Iberia, Southwest France

DF27 R1b1a1a2a1a2a [5]

M153 [R1b1b2a2c] R1b1a1a2a1a2a1a1a1? [5a] Basque, Gascon, Iberia

M65 Basque

DF27 / M167 / SRY2627 [R1b1b2a2d] R1b1a1a2a1a2a1b1a1 [5b] Basque, (especially) Catalonia, Cornwall, Wales, Northeast Spain, Portugal, Southwest France, Bavaria, Dutch, Belgians

M126 [R1b1b2a2g1] [6a] Alpine Celts

M160 [R1b1b2a2g] [6b] Alpine Celts

U152 / S28 [R1b1b2a2g] R1b1a1a2a1a2b [6] [southern] Switzerland, Central/Eastern France and Italy [N & C], Jutland, Southwestern Norway, [R1b1b2h1?]

S68 [R1b1b2a2h] Sweden, Scotland

L165 / S28 Scotland

L2/ S139 Italy, Germany, Belgium, Britain, Ireland, Norway 

L20 / S144 England, France, Italy

M529 / L21 / S145 R1b1a1a2a1a2c1[7] Brittany and Ireland, Scotland and Wales, Northwest France, Southwest Norway 

P66 [R1b1b2a2f] Northern Italy

M37 [R1b1b2a2a] Ireland

L193 Scottish Borders 

M222 [DF49/DF23] [R1b1b2a2e/R1b1a2a1a1b4b] R1b1a1a2a1a2c1a? [7a] West Scotland, Scots-Irish, North West Ireland

L238 / S182 Norway, Sweden 

DF19 Germany, Benelux, Britain

DF1 / L513 / S215 / L193 / S176 Scotland

L226 / S168 Central and Western Ireland

L159 / S169 Isle of Man, Hebrides, Norway, Denmark, Northern France, England, Scotland, Ireland

L176 / S179 Northeast Spain, Southwest France

L165 / S68 [R1b1b2a2h] Scandinavia, England, Northern Isles and Outer Hebrides

L1335 Scotland [Pict]

L371 Wales

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